tRNA Cys-GCA-4-1中数字4和1代表了什么含义？...

It is likely an id for tRNA Cys-GCA non-coding RNA gene. In HUGO Gene Nomenclature Committee (HGNC) data base, tRNA Cys-GCA non-coding RNA list 32 different genes with different 2 digital number as gene id:

http://www.genenames.org/cgi-bin/genefamilies/set/842